Emotion is a central dimension of spoken communication, yet, we still lack a mechanistic account of how modern large audio-language models (LALMs) encode it internally. We present the first neuron-level interpretability study of emotion-sensitive neurons (ESNs) in LALMs and provide causal evidence supporting the existence of such units in Qwen2.5-Omni, Kimi-Audio, and Audio Flamingo 3. Across these three widely used open-source models, we compare frequency-, entropy-, mean-deviation-, and contrast-based neuron selectors on multiple emotion recognition benchmarks. Using inference-time interventions, we reveal a consistent emotion-specific signature: deactivating neurons selected for a given emotion disproportionately degrades recognition of that emotion while largely preserving other classes, whereas targeted steering amplifies these units to bias predictions toward the target emotion. These effects arise with modest amounts of identification data and scale systematically with intervention strength. We further observe that ESNs exhibit non-uniform layer-wise clustering with partial cross-dataset transfer. Taken together, our results offer a causal, neuron-level account of emotion decisions in LALMs and highlight targeted neuron interventions as an actionable handle for controllable affective behaviors.

Conversational Speech Synthesis (CSS) aims to generate speech with natural prosody by understanding the multimodal dialogue history (MDH). The latest work predicts the accurate prosody expression of the target utterance by modeling the utterance-level interaction characteristics of MDH and the target utterance. However, MDH contains fine-grained semantic and prosody knowledge at the word level. Existing methods overlook the fine-grained semantic and prosodic interaction modeling. To address this gap, we propose MFCIG-CSS, a novel Multimodal Fine-grained Context Interaction Graph-based CSS system. Our approach constructs two specialized multimodal fine-grained dialogue interaction graphs: a semantic interaction graph and a prosody interaction graph. These two interaction graphs effectively encode interactions between word-level semantics, prosody, and their influence on subsequent utterances in MDH. The encoded interaction features are then leveraged to enhance synthesized speech with natural conversational prosody. Experiments on the DailyTalk dataset demonstrate that MFCIG-CSS outperforms all baseline models in terms of prosodic expressiveness. Code and speech samples are available at https://github.com/AI-S2-Lab/MFCIG-CSS.